《Table 1.Details on the concatenated alignment of the four target regions used in this study.》

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《Aedeagus evolution promotes speciation A primary pattern in rove beetle phylogeny》

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aThree codons at the 5′end of COI excluded due to gene length polymorphism bThirty positions of Wingless excluded due to ambiguous alignment cOne codon at the middle of Tp excluded due to ambiguous alignment dPositions excluded due to ambiguous alignment

Phylogenetic analyses were based on one nuclear ribosomal（28S=899–1132 bp）and three protein coding（COI=792–834 bp，Wg=366–438 bp，Tp=600–726 bp）loci.The concatenated sequence alignment contained 61 sequences and 3085positions after trimming（Table 1）.The likelihood models identified by Modeltest（AIC）were given in Table 2.In the Bayesian analysis（Fig.1），the subfamily Staphylininae was recovered as monophyletic with a high posterior probability support value（pp=0.99）.The tribe Platyprosopini was in the basal of the subfamily Staphylininae.Excluding the tribe Platyprosopini，another five tribes made up two main clades:one clade including the tribes Staphylinini and Arrowinini（Clade A in Fig.1），the other clade containing Xantholinini，Othiini and Diochini three tribes（Clade B in Fig.1）.In Clade A，several small clades were all with a 1.0 posterior probability support value，including the clades such as Staphylinina+Anisolinina，Xanthopygina，Philonthina and Tanygnathina+Hyptiomina.In Clade B，Xantholinini and Othiini two subclades were also well supported（pp=1.0）.The maximum likelihood analysis generated a similar topology as Bayesian analysis except for the placement of the subtribe Amblyopinina and the genus Algon（Supplementary Fig.S3）.