《Table 3.Summary of the Beast analysis for the age and nuclear substitution rates of important nodes

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《Aedeagus evolution promotes speciation A primary pattern in rove beetle phylogeny》

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*HPD represents the highest posterior density

According to the relaxed molecular clock analysis of the combined data，the origin of the subfamily Staphylininae was estimated at about 176.11 million years ago（Mya）with a 95%confidence interval from 161.38 Mya to 190.78 Mya.The Beast analysis for the age of important clades，such as Clade A and Clade B，and clades Staphylinina+Anisolinina，Xanthopygina，Philonthina，Tanygnathina+Hyptiomina，Othiini and Xantholinini，were shown in Table 3.Here we selected the Bayesian tree to show the phylogenetic pattern of aedeagus morphology and divergence time estimation（Fig.2）.Morphological analysis revealed mainly three types of aedeagus structural variations:（1）aedeagus with a single paramere（Asp）；（2）aedeagus with a bifurcated paramere（Abp）；and（3）aedeagus with a pair of parameres（App）.