《Table 2–List of species with assembled pan-genomes.》

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《The role of genomic structural variation in the genetic improvement of polyploid crops》

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n.a.，data not available/presented.

All three major types of genome structural variation（CNV，PAV，and HE）have been linked to important agronomic traits in polyploid crops（Table 1），despite the fact that genomic resources are still sparse for many of these species.The production of multiple reference genomes per species and the subsequent construction of“pan-genomes”that include all（or almost all）DNA present in all individuals within a species have been under way for only a short time，and most of our important crop species are still lacking pan-genome resources（Table 2）.Fortunately，these resources are becoming increasingly available with the advent of new sequencing technologies，and with the corresponding drop in price and time required for genome sequencing.In the short term，detection of genome structural variation at lower costs can be optimized by combining multiple technologies，such as short-read sequencing with longer reads or optical mapping，or by the use of older methods such as molecular cytogenetics or genotyping arrays[50，66].