《Table 1–ANOVA of disease severity in the BC1F2:3population.》

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《Identification of stably expressed QTL for resistance to black shank disease in tobacco(Nicotiana tabacum L.) line Beinhart 1000-1》

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df，degrees of freedom.

The BS responses of the parents were significantly different；BH expressed a very high level of resistance with a score of 0for both disease severity（DS）and disease index（DI），whereas XHJ was highly susceptible with of DS 9 and DI of 84.50%–100.00%for different environments（Fig.2）.The average value of DS for the F2population was 3.20，and for the BC1F2，3.57.For the BC1F2:3population，the mean DI for the field stage and seedling stage tests were 42.02%and 41.86%，respectively.All populations displayed significantly skewed distributions in the direction of resistance，indicating that both major genes and polygenes controlled resistance to BS.As shown in Table 1，the ANOVA detected highly significant differences among lines（P<0.001）and no significant differences between environments and lines（P=0.289）.